Update README.md

Author: davidborland

README.md

@@ -5,4 +5,6 @@ See the [wiki](https://github.com/RENCI/Segmentor/wiki/) for more information on
 
 Visit the [nuclei ninja](https://www.nucleininja.org/) website to find out more about the NIH-funded project driving the development of this tool.
 
+See the [paper](https://bmcbioinformatics.biomedcentral.com/articles/10.1186/s12859-021-04202-8) on Segmentor published in BMC Bioinformatics.
+
 ![image](https://user-images.githubusercontent.com/289957/93640079-f906c500-f9c7-11ea-81b6-d9fdbfb5f479.png)